I think Zelena may sacrifice her father’s heart too or maybe her adoptive mother’s. There’s a lot of parallels going on with Regina and Zelena yet still being different.
What about the heart of a former lover?
Regina started down the path toward the curse because of Daniel’s death–his heart being crushed by Cora. What if Zelena crushed the heart of her own lover?
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"He was a lot of things to me" "The only conclusion was love"